Basidiomata annual, centrally to laterally stipitate, coriaceous to corky. Pileus single, suborbicular to umbelliform, up to 8.5 cm diam and 5 mm thick. Pileal surface yellow to yellowish brown when dry, dull, tomentose, with concentric furrows and radial wrinkles, sagging at the centre; margin acute to obtuse, entire, wavy and incurved when dry. Pore surface white to cream buff when fresh, colour unchanging when bruised, pale yellow when dry, even ferruginous in old specimens; pores circular to angular, 6–7 per mm; dissepiments slightly thick, entire. Context white to cream, without dark resinous lines, corky, up to 2 mm thick. Tubes concolorous with pore surface, hard corky, up to 3 mm long. Stipe concolorous with pileal surface, cylindrical and hollow, swollen at base, up to 7.5 cm long and 1 cm diam. Hyphal system trimitic; generative hyphae with clamp connections, all hyphae IKI–, CB+; tissues darkening in KOH. Generative hyphae in context colourless, thin-walled, 2–4 μm diam; skeletal hyphae in context colourless to pale yellow, thick-walled with a wide to narrow lumen or subsolid, arboriform branched and flexuous, 3–6 μm diam; binding hyphae in context colourless, subsolid, branched and flexuous, 1–2 μm diam. Generative hyphae in tubes colourless, thin-walled, 2–3 μm diam; skeletal hyphae in tubes colourless to pale yellow, thick-walled with a wide to narrow lumen or subsolid, arboriform branched and flexuous, 2–5 μm diam; binding hyphae in tubes colourless, subsolid, branched and flexuous, 1–2 μm diam. Pileal cover composed of clamped generative hyphae, thin- to thick-walled, apical cells clavate and inflated, often slanting to one side, yellowish brown, about 60–80 × 5–10 μm, forming a regular palisade. Cystidia absent; cystidioles clavate, colourless, thin-walled, 18–20 × 4–5 μm. Basidia barrel-shaped to clavate, colourless, thin-walled, 13–23 × 11–14 μm; basidioles in shape similar to basidia, colourless, thin-walled, 13–17 × 6–10 μm. Basidiospores globose to subglobose, pale yellow, IKI–, CB+, with double and distinctly thick walls, exospore wall smooth, endospore wall with conspicuous spinules, (7–)7.2– 8.5(–8.7) × (5.8–)6.7–8(–8.3) μm, L = 7.87 μm, W = 7.37 μm, Q = 1.06–1.07 (n = 60/2). Under SEM, exospore wall uneven or foveolate; endospore wall with some hollow and columnar spinules which persist to exospore wall forming holes.
Notes — Amauroderma austrosinense was described from the tropical part of China (Zhao et al. 1984). It has the typical features such as white to straw colour pore surface when fresh and colour unchanging when bruised, cream context, hollow and columnar spinules on endospore wall which persist to the exospore wall forming the holes (Fig. 3a–b) that characterise Foraminispora (Costa-Rezende et al. 2017). In the phylogenetic analyses, A. austrosinense was shown as a distinct lineage in Foraminispora with high support (100 % ML, 1.00 BPP). Therefore, we transferred A. austrosinense to Foraminispora as a new combination.
华南假芝 图93
Amauroderma austrosinense J.D. Zhao et L.W. Hsu, Acta Mycol. Sinica 3(1):20,Fig.5,Pl.1-3,1984; Zhao, The Ganodermataceae in China (Chinese edition). p. 211，Fig.78，1989.
担子果一年生，有柄，伞状，木质。菌盖近圆形或稍呈不规则形，直径3.5-4cm，厚 3-4mm，中央稍平坦，表面呈可可褐色或淡黄褐色，具较稠密的深褐色同心环纹，有不规则的放射状沟;边缘钝，略向内卷，有时波状;菌肉呈淡白色，厚2-3mm。菌管与菌肉同色，长1-3mm;孔面淡白色，后变淡褐色;管口略圆形，完整每毫米5-6个;菌柄近中生、偏生或侧生，与菌盖同色，中空，肉淡白色，长4--12cm，粗0.5-1cm，粗细不均匀，有时下部变扁。
皮壳构造由近平行的生殖菌丝构成，彼此胶粘在一起不易分离开，呈淡红褐色，其组成菌丝的顶端膨大或否，有的顶端稍尖细，下部有锁状联合，其间杂有淡褐色厚壁菌丝，皮壳表面有游离的透明生殖菌丝，似与毛皮壳型类似。
菌丝系统三体型:生殖菌丝无色透明，薄壁，有横膈膜，直径3.5-4μm;骨架菌丝淡黄色或几乎无色，具树状分枝;骨架干直径3-4μm，分枝末端形成鞭毛状无色缠绕菌丝;缠绕菌丝无色，厚壁，分枝，弯曲，直径1.5-2μm。
担孢子近球形或球形，双层壁，外壁无色透明，平滑，内壁淡黄色，具稠密的小刺，7.5-9x6.5-8.3μm。
模式产地:中国(海南)。
生境:生于林中地上。
研究标本:海南坝王岭雅加林场(42695模式):广西大瑶山(42790)
世界分布:中国。
讨论:此种的特点是担子果呈伞状，菌盖有较稠密的同心环沟，具有不规则的放射状沟，担孢子近球形，7.5-9x6.5-8.3μm。尚未见其近似种。
